
Masumoto, K., V. Novak, K. Akita, C.-F. Lee. 2019. A revisional study of the subfamily Alleculinae (Coleoptera: Tenebrionidae) of Taiwan (Part 4). Ten new species of genera Allecula, Borbonalia, Borboresthes, Pseudohymenalia and Upinella. Folia Heyrovskyana, series A, 27: 20-51.